After watching the Wales v England game, I'd say Gethin Jenkins is up there with the fittest in union, considering the 18+ stone he's carrying around. Back in 2005 I remember a comment in one of the games that Gethin had the best bleep test results in the squad, and he seems even fitter today! putting in 17 tackles, grappling for the ball ruck time, clearing out umpteen rucks, supporting players for offloads etc. All this on top of a props usual duties of scrummaging. I can't emagine many people could keep that up for 80min's let alone carrying the weight around he does. On ScrumV on Sun night, Kingsley Jones commented that back in Bridgend days, when Gethin was just coming through as a fat teenager he was beating them all on cross country runs. Some people are naturals I suppose.
I agree with the majority on here, I doubt League conditioning could be improved much, for what they do at the moment, they can easily last the 80mins required without dropping much in tempo and tenacity, same for Union. Union has deffinatly caught up though, no longer do you find overwight players (well Dunning and Adam Jones aside, though Jones seems much better) up front, all are well conditioned etc.